Skip to content
Atlas of Computational Cell Reprogramming

← Methods

L3 · Explicit model-based inverse intervention D P

CellOracle

Kamimoto K, Stringa B, Hoffmann CM, Jindal K, Solnica-Krezel L, Morris SA

2023 · Nature

Cell identity is governed by the complex regulation of gene expression, represented as gene-regulatory networks1.

Abstract

From the original paper, Nature · PubMed

Cell identity is governed by the complex regulation of gene expression, represented as gene-regulatory networks1. Here we use gene-regulatory networks inferred from single-cell multi-omics data to perform in silico transcription factor perturbations, simulating the consequent changes in cell identity using only unperturbed wild-type data. We apply this machine-learning-based approach, CellOracle, to well-established paradigms-mouse and human haematopoiesis, and zebrafish embryogenesis-and we correctly model reported changes in phenotype that occur as a result of transcription factor perturbation. Through systematic in silico transcription factor perturbation in the developing zebrafish, we simulate and experimentally validate a previously unreported phenotype that results from the loss of noto, an established notochord regulator. Furthermore, we identify an axial mesoderm regulator, lhx1a. Together, these results show that CellOracle can be used to analyse the regulation of cell identity by transcription factors, and can provide mechanistic insights into development and differentiation.

Summary

Editorial summary pending review by the maintainer. The paper's own abstract appears above; the Atlas summary in the maintainer's voice will explain how CellOracle relates to the cross-modality inverse-design framework of the review.

Why this level

Level 3 because candidate interventions enter an explicit forward operator FθuF_{\theta_u} and the predicted post-intervention outcome is what scores each candidate. Representation family is executable intervention model. Cited in §3.4 of the review. Editorial rationale pending review by the maintainer.

Classification

Level
L3
Representation
Executable intervention model
Modalities
D, P
Intervention
Transcription factors
Framework
Boolean network

Software

Reproducibility
4 of 4
FAIR4RS
3 of 5

Last audited 2026-05-24

Citation

Kamimoto K et al. (2023). Dissecting cell identity via network inference and in silico gene perturbation., Nature.

DOI: 10.1038/s41586-022-05688-9

PMID: 36755098

BibTeX
@article{celloracle2023,
  title  = {Dissecting cell identity via network inference and in silico gene perturbation.},
  author = {Kamimoto K et al.},
  year   = {2023},
  journal = {Nature},
  pmid = {36755098},
  doi  = {10.1038/s41586-022-05688-9}
}